Jonathan, the world's oldest living land animal, died today at an estimated age of 193.
Jonathan was a Seychelles giant tortoise (Aldabrachelys gigantea hololissa), a subspecies of the Aldabra giant tortoise (Aldabrachelys gigantea), but he spent most of his life on the remote South Atlantic island of St Helena, where he was brought when he was about 50.
Jonathan (left) is pictured with another giant tortoise in 1886, when they were aged 53–54.
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